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Semiochemicals of Ovophis okinavensis, the Ryukyu Island pit viper

Phylum:  Chordata
Subphylum:  Vertebrata
Class:  Reptilia
Order:  Squamata
Family:  Viperidae
Subfamily:  Crotalinae
Genus:  Ovophis
Tribe:  
Author:  Boulenger
�
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Image: Google
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Biology: Google
Distribution: Google
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Bedoukain RussellIPM

Semiochemical(s):

  Villar-Briones A  2018  Toxins  10: 392   
    citric acid    �Relative ratio of the component�   �(*) indicates that compound is active�   P
�Category of the chemical signal
A -   Attractant
Al -   Allomone
K -   Kairomone
P -   Pheromone
Sy -   Synomone�
  
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		Okinawa, Japan 
    VPPDH    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    pEKW    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    pENW    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    pEGGPR    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    pEWQ    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    WPPGPPIPP    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    pE(NH)    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    VPPGPPGPPP    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    PGPGPPP    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    pETAP    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    PP    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    2-oxoarginine    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    itaconic acid    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    pEQ    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    creatine    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    cytidine    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    pEE    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    choline    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    4-guanidinobutyric acid    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    imidazoleacetic acid    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    pEKS    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    pEK    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    pEH    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    pEG    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    pEGE    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    non-pE pro peptide    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    l-proline    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    4-oxoglutaramate    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    pE peptide    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    aconitic acid    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    pER    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    pEEGT    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    betaine    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    glutarimide    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    oxoglutaric acid    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    l-carnitine    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    pseudouridine    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    taurine    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    stearic acid    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    pEGRP    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    AP    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    3-imidazol-5-yl-pyruvate    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    hydroxyphenylacetic acid    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    quinolin-8-ol    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    creatinine    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    3-1-indol-3-yl-acrylic acid    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    palmitic acid    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    l-pyroglutamate    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    prolylleucine    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    choline phosphate    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    pES    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    phenylpyruvic acid    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    acetylcholine    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    lauric acid    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    IP    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    pESN    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    p-cresol    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    4-hydroxyphenylpyruvic acid    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    hypoxanthine    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    4-hydroxybenzaldehyde    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    pEND    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    glutamine    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    pipecolic acid    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    succinic acid    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    carnosine    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    N-acetylneuraminic acid    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    l-malic acid    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    uric acid    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    pEN    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    glycerophosphocholine    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    12-hydroxystearic acid    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    VP    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    oleic acid    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    1H-indole    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    adenosine    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    inosine    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    l-isoleucine    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    l-glutamic acid    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    pETGG    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    pENWPAPK    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    4-guanidinobutanal    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    pelargonic acid    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    l-threonine    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    pEEWPPGHHIPP    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    l-arginine    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    N-palmitoylsphingomyelin    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
 
  Yamazaki Y  2004  Toxicon  44: 227   
    okinavin    �Relative ratio of the component�   �(*) indicates that compound is active�   P
�Category of the chemical signal
A -   Attractant
Al -   Allomone
K -   Kairomone
P -   Pheromone
Sy -   Synomone�
  
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		Okinawa, Japan 
 
  Tan NH  1989  Comp. Biochem. Physiol. B  93: 757   
    hyaluronidase    �Relative ratio of the component�   �(*) indicates that compound is active�   P
�Category of the chemical signal
A -   Attractant
Al -   Allomone
K -   Kairomone
P -   Pheromone
Sy -   Synomone�
  
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		USA 
    protease    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    5'-nucleotidase    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    arginine esterase    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    LAAO    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    alkaline phosphatase    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    PDE    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    PLA2    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
 
  Mebs D  1970  Int. J. Biochem.  2: 335   
    PDE    �Relative ratio of the component�   �(*) indicates that compound is active�   P
�Category of the chemical signal
A -   Attractant
Al -   Allomone
K -   Kairomone
P -   Pheromone
Sy -   Synomone�
  
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		Miami, Florida, USA 
    5'-adenylic acid    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    BAEE    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    LAAO    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
    caseinase    �Relative ratio of the component�   �(*) indicates that compound is active�    
�Source of the chemical signal
F -   Female
H -   Host (could be of plant or animal origin)
L -   Lure
M -   Male
M&F -   Male and Female
S -   Solider
Q -   Queen
W -   Worker�
  
�Amount
pg -   Picogram
ng -   Nanogram
�g -   Microgram
mg -   Milligram
g -   Gram�
   		 
 

Reference(s):

Villar-Briones, A., and Aird, S.D. 2018. Organic and peptidyl constituents of snake venoms: The picture is vastly more complex than we imagined. Toxins. 10:392-441.
 
Yamazaki, Y., and Morita, T. 2004. Structure and function of snake venom cysteine-rich secretory proteins. Toxicon. 44:227-231.
 
Tan, N.-H., Armugam, A., and Tan, C.-S. 1989. A comparative study of the enzymatic and toxic properties of venoms of the Asian lance-headed pit viper (genus Trimeresurus). Comp. Biochem. Physiol. B. 93:757-762.
 
Mebs, D. 1970. A comparative study of enzyme activities in snake venoms. Int. J. Biochem. 2:335-342.
 
 
Citation: El-Sayed AM 2025. The Pherobase: Database of Pheromones and Semiochemicals. <http://www.pherobase.com>.
Ⓒ 2003-2025 The Pherobase - Extensive Database of Pheromones and Semiochemicals. Ashraf M. El-Sayed.
Page created on 19-January-2025